He's a Dead Apostle that uses his bee familiars to increase his number of Ghouls.

The rulers implanted tiny machines into the slaves' heads to control them.

...can serve only a righteous human.

was his exploitation as a Guardian, even after his death.

I'm the one who used that! What nonsense are you talking...?!

Even though they be fake, I cannot admire those that use living things so thoughtlessly...

And if the rebels have themselves a dragon, General Tullius is the only one who can stop them.

Some transform into giant bats, others communicate with animals, command them.

Summon and tame the djinn. It's not likely to be happy. Stand at the ready.

His books strongly suggest he did. He describes taming one, harnessing energy from its power.